function [ihc_out, state] = lyon2011_ihcstep(bm_out, coeffs, state)
%LYON2011_IHCSTEP update of inner-hair-cell (IHC) model
%
% Usage: [ihc_out, state] = lyon2011_ihcstep(bm_out, coeffs, state);
%
% TODO(mvelimirovic): rename coeffs parameter to IHC_coeffs.
% LYON2011_IHCSTEP updates the inner-hair-cell (IHC) model, including the
% detection nonlinearity and one or two capacitor state variables. This
% file is part of an implementation of Lyon's cochlear model:
% "Cascade of Asymmetric Resonators with Fast-Acting Compression"
%
%
% See also: lyon2011 demo_lyon2011
%
% References:
% R. F. Lyon. Cascades of two-pole–two-zero asymmetric resonators are
% good models of peripheral auditory function. J. Acoust. Soc. Am.,
% 130(6), 2011.
%
%
% Url: http://amtoolbox.org/amt-1.5.0/doc/modelstages/lyon2011_ihcstep.php
% #StatusDoc: Good
% #StatusCode: Good
% #Verification: Unknown
% #License: Apache2
% #Author: Richard F. Lyon (2013): original implementation (https://github.com/google/carfac)
% #Author: Amin Saremi (2016): adaptations for the AMT
% #Author: Clara Hollomey (2021): integration in the AMT 1.0
% #Author: Richard Lyon (2022): bug fixes for AMT
% #Author: Mihajlo Velimirovic (2022): implementation of the option ihc_potential
% This file is licensed unter the Apache License Version 2.0 which details can
% be found in the AMT directory "licences" and at
% <http://www.apache.org/licenses/LICENSE-2.0>.
% You must not use this file except in compliance with the Apache License
% Version 2.0. Unless required by applicable law or agreed to in writing, this
% file is distributed on an "as is" basis, without warranties or conditions
% of any kind, either express or implied.
% #Author: Amin Saremi (2016) adaptations for the AMT (based on
% <https://github.com/google/carfac>, Richard F. Lyon)
% #Author: Richard Lyon (2023) further adaptations and new two-cap model.
% If ac_coeff is positive, AC couple the bm_out, with 20 Hz corner.
% If ac_coeff is 0 here, it is done in CAR step.
if coeffs.ac_coeff > 0
ac_diff = bm_out - state.ac_coupler;
state.ac_coupler = state.ac_coupler + coeffs.ac_coeff * ac_diff;
else
ac_diff = bm_out;
end
if coeffs.just_hwr
ihc_out = min(2, max(0, ac_diff)); % limit it for stability
else
conductance = lyon2011_detect(ac_diff); % rectifying nonlinearity
if coeffs.one_cap;
% Output comes from receptor current like in Hall and Allen's models.
ihc_out = conductance .* state.cap_voltage;
state.cap_voltage = state.cap_voltage - ...
ihc_out .* coeffs.out_rate + ...
(1 - state.cap_voltage) .* coeffs.in_rate;
ihc_out = ihc_out * coeffs.output_gain;
% Smooth it twice with LPF:
state.lpf1_state = state.lpf1_state + coeffs.lpf_coeff * ...
(ihc_out - state.lpf1_state);
state.lpf2_state = state.lpf2_state + coeffs.lpf_coeff * ...
(state.lpf1_state - state.lpf2_state);
ihc_out = state.lpf2_state - coeffs.rest_output;
else
% Change to 2-cap version mediated by receptor potential at cap1:
% Geisler book fig 8.4 suggests 40 to 800 Hz corner.
receptor_current = conductance .* state.cap1_voltage;
% "out" means charge depletion; "in" means restoration toward 1.
state.cap1_voltage = state.cap1_voltage - ...
receptor_current .* coeffs.out1_rate + ...
(1 - state.cap1_voltage) .* coeffs.in1_rate;
% Amount of depletion below 1 is receptor potential.
receptor_potential = 1 - state.cap1_voltage; % Already smooth.
% Identity map from receptor potential to neurotransmitter conductance.
ihc_out = receptor_potential .* state.cap2_voltage; % Now a current.
% cap2 represents transmitter store; another adaptive gain.
% Deplete the transmitter store like in Meddis models:
state.cap2_voltage = state.cap2_voltage - ...
ihc_out .* coeffs.out2_rate + ...
(1 - state.cap2_voltage) .* coeffs.in2_rate;
% Awkwardly, gain needs to be here for the init states to be right.
ihc_out = ihc_out * coeffs.output_gain;
% smooth once more with LPF (receptor potential was already smooth):
state.lpf1_state = state.lpf1_state + coeffs.lpf_coeff * ...
(ihc_out - state.lpf1_state);
ihc_out = state.lpf1_state - coeffs.rest_output;
end
end
state.ihc_accum = state.ihc_accum + ihc_out; % for where decimated output is useful